New additions are indicated by an asterisk (*). Last update: 28 February 2016.

Kindly send suggestions of HAB links you would like us to add, corrections for broken links, or corrections to the text that describes the link, to Karen Lebret.


* Burrell, Stephen. 2015. Reference Material Development for Paralytic Shell sh Poisoning Toxins and Associated Analytical Applications. PhD Thesis. Dublin Institute of Technology, Dublin, Ireland, 284 p. Link to PDF.

Sassenhagen, Ingrid. 2015. Physical and biological dispersal barriers in invasive, bloom-forming microalgae. PhD Thesis. Lund University. Lund, Sweden. Link to PDF.


Lim, Hong Chang. 2014. Morphology, taxonomy and genetics of tropical diatom Pseudo-nitzschia species (Bacillariophyceae) in Malaysian waters. PhD Thesis. Faculty of Resource Science and Technology, Universiti Malaysia Sarawak, Malaysia, 180 p. PDF file in ResearchGate; 7 MB

Zastepa, Arthur. 2014. Fate and Persistence of Microcystin Congeners in Lakes and Lake Sediments. PhD thesis. University of Ottawa.


Bouchouicha, Donia. 2013. Dynamique saisonnières des microalgues nuisibles dans la lagune de Bizerte et identification d’une nouvelle espèce Nitzschia bizertensis sp. nov. productrice d’acide domoïque. Mémoire de thèse, Faculté des Sciences de Bizerte - Université de Carthage. 231 p.


Bakke, Marit Jørgensen. 2012. Effects of algal-produced neurotoxins on fish. PhD thesis. Norwegian School of Veterinary Science, 7 June 2012. Information

Guannel, Michele L. 2012. Marine microbial biogeography from microscopic to global scales: distributions of the diatom Pseudo-nitzschia and co-existing bacteria. PhD thesis, University of Washington. Link to abstract

Henningsen, Brooks M. 2012. Bioinformatic and gene expression analysis of an SLC6 homolog in the toxin-producing marine diatom Pseudo-nitzschia multiseries. MS thesis. Plymouth State University, Biological Sciences, 218 p.

Lebret, Karen. 2012. A Phytoplankton Invasion: Population Genetics, Phylogeography, and Invasion Success of Gonyostomum semen. 6 December 2012. Doctoral Dissertation. Lund University. Lund, Sweden. Link to PDF

Liefer, Justin D. 2012. Phytoplankton community ctructure and the physiological ecology of the toxic bloom-forming diatom Pseudo-nitzschia spp. in coastal Alabama. Ph.D. Dissertation, University of South Alabama, Mobile, AL, May 2012, 355 p. ProQuest abstract and link to preview PDF file.

O’dea, Sheila. 2012. Occurrence, toxicity, and diversity of Pseudo-nitzschia in Florida coastal waters. Master of Science Thesis, University of South Florida, 103 p. Download PDF file from Scholar Commons; 2.2 MB.

Timmerman, Amanda. 2012. Hidden thin layers of toxic diatoms in a coastal bay. Master of Science in Oceanography. University of Hawaii at Mānoa. Abstract


Su, Suriyanti Nyun-Pau. 2011. Ecology and taxonomy of amnesic shellfish poisoning (ASP) diatom Pseudo-nitzschia species (Bacillariophyceae) in Malaysian waters. M.Sc. Dissertation, Faculty of Resource Science and Technology, Universiti Malaysia Sarawak, Malaysia. 134 pp.

Lim, Hong Chang. 2011. Morphological and molecular characterization of potential amnesic shellfish poisoning toxin-producing diatom Pseudo-nitzschia (Bacillariophyceae) from Malaysia. Master of Science Thesis, Universiti Malaysia Sarawak, Sarawak, Malaysia, 138 p. Access to full text via ResearchGate (1.8 MB)

Giussani, Valentina. 2011. Dynamics of dinoflagellate Ostreopsis ovata bloom along Genoa coast during the summer of 2010 and toxic effects of microalga on invertebrate and vertebrate marine organisms. University of Genoa. Italy.

Gracia, Stéphanie. 2011. Distribution spatiale des kystes benthiques d’Alexandrium tamarense suite à une floraison massive dans l’estuaire du Saint-Laurent. MSc Thesis, Université du Québec à Rimouski, Québec, Canada. 21 December 2011.

Calu, Guillaume. 2011. Contribution à l'étude de la toxicité de Pseudo-nitzschia: régulation de la production d'acide domoïque et synthèse d'analogues chimiques. PhD Thesis, Université de Nantes, Nantes, France, 212 p. Link to announcement of thesis defence (5 July 2011)

Santiago-Morales, Ivonne Sandra. 2011. Potencial tóxico de cepas de Pseudo-nitzschia aisladas de la costa occidental de Baja California. [Toxic potential of Pseudo-nitzschia strains isolated from the west coast of Baja California]. PhD Thesis. Centro de Investigación Científica y de Educación Superior de Ensenada. Ensenada, Baja California, Mexico. September 14. 161 p. Link to PDF file


Cabrales Talavera, Guadalupe. 2010. Distribución espacial y temporal de diatomeas del género Pseudo-nitzschia spp y la concentración de ácido domoico en la región de la Bahía de Todos Santos, B.C., México. Master's Thesis. Centro de Investigación Científica y de Educación Superior de Ensenada. Ensenada, Baja California, Mexico. Information on thesis

Hubbard, Katherine Ada. 2010.Ecological investigations of genetic diversity in the diatom genus Pseudo-nitzschia. PhD Thesis, University of Washington, Seattle, WA, 302 p.

Nguyen, Dieu. 2010. Asymmetric allylation of aldehydes catalyzed by formamide organocatalysts. Studies towards the synthesis of L- domoic acid side chain. PhD Thesis, University of South Carolina. Abstract

Sahraoui, Inès. 2010. Écologie, diversité et toxicité des diatomées du genre Pseudo-nitzschia H. Peragallo dans la lagune de Bizerte. PhD Thesis, Université 7 Novembre à Carthage, Bizerte, Tunisia, 267 p.

Sedehizadeh, Simon. 2010. Towards the total synthesis of domoic acid and the isodomoic acids. PhD Thesis, The University of Manchester, UK, 265 p. (PDF file; 2.6 MB)

Tesson, Sylvie V.M. 2010. Population genetic structure of a planktonic diatom in the Gulf of Naples: Pseudo-nitzschia multistriata. PhD Thesis, October 2010. The Open University of London. 285 p. Abstract

Varkitzi, I. 2010. Ecology of harmful microalgae and impact on the marine benthic ecosystem. PhD Thesis. University of Athens, Greece, 347 p.

Yap-Dejeto, Leni G. 2010. Development of practical quantification protocols of the marine diatom Pseudo-nitzschia spp. for ecological studies. PhD Thesis, March 2010. The University of Tokyo. 176 p.


Casteleyn, Griet. 2009. Species structure and biogeography of Pseudo-nitzschia pungens. PhD thesis. Laboratory of Protistology & Aquatic Ecology, Biology Department, Ghent University, Ghent, Belgium. 201 p. Information

Del Rio, Ross P. 2009. Neurotoxin in a Louisiana estuary: quantitative analysis of domoic acid in gulf menhaden (Brevoortia patronus) and qualitative modeling of links in a shark nursery. MSc Tesis. Louisiana State University and Agricultural and Mechanical College, Baton Rouge, Lousiana, USA. ix + 102 p. Abstract and link to PDF file

Kassab, Refaie Mohammad Refaie. 2009. Nickel-catalyzed carbon-carbon bond forming reactions involving nickel-carbene insertion and studies toward the total synthesis of domoic acid. PhD Thesis, AAT 3371706, Wayne State University, 277 p. Abstract and link to "View More", including a preview of first part of thesis

Mafra, Luiz L., Jr. 2009. Comparative domoic acid kinetics in oysters (Crassostrea virginica) and mussels (Mytilus edulis) feeding on Pseudo-nitzschia multiseries: implications for species-specific management. PhD Thesis, Dalhousie University, Halifax, NS, Canada, 233 p. Abstract

Moustafa, A. 2009. Evolutionary and functional genomics of photosynthetic eukaryotes. PhD Thesis, University of Iowa, USA. Abstract.


Moccia, Lauren Paul. 2008. Cloning and characterization of a novel ferratin from the marine diatom Pseudo-nitzschia multiseries. M.Sc. Thesis. University of British Columbia, Vancouver, BC, Canada, 87 p. PDF file; 3.15 MB

D'Alelio, Domenico. 2008. The life cycle of the diatom Pseudo-nitzschia multistriata: experimental and in situ approaches. PhD Thesis. University of Messina, Italy, 172 p.


Anderson, Clarissa R. 2007. Environmental controls of phytoplankton community structure in the Santa Barbara Channel, California: application to the dynamics and detection of toxigenic Pseudo-nitzschia blooms. PhD Thesis, University of California, Santa Barbara, 166 p. Abstract

Thessen, Anne E. 2007. Taxonomy and ecophysiology of Pseudo-nitzschia in the Chesapeake Bay. PhD thesis, University of Maryland, College Park, MD, USA, 231 p. PDF (2.1 MB)

Vranyac-Tramoundanas, Alexandra. 2007. Domoic acid-induced cardiac damage: an in vitro and in vivo investigation. PhD Thesis, University of Otago, New Zealand. Abstract


Adams, Nicolaus G. 2006. Genetic differentiation of Pseudo-nitzschia pungens from the Pacific Northwest and the North Sea. Master’s Thesis, University of Washington, Seattle, 96 p. PDF (1.9 MB)

Mengelt, Claudia. 2006. Ultraviolet photoecology, dark survival, and seasonal abundance of Pseudo-nitzschia australis and Pseudo-nitzschia multiseries in coastal waters of central California. PhD Thesis, University of California, Santa Barbara, 208 p. Abstract and link to PDF file

Olson, Michael Brady. 2006. On the population ecology of the toxigenic diatom genus Pseudo-nitzschia: perspectives from the growth and mortality environments. PhD Thesis, University of Washington, Seattle, WA, USA, 160 p. (August 30, 2006)


O'Higgins, L. 2005. An investigation of domoic acid producing diatoms (Bacillariophyceae) from coastal and shelf waters of Northwest Europe. PhD thesis, National University of Ireland, Galway, 199 p.

Lips, Inga. 2005. Abiotic factors controlling the cyanobacterial bloom occurrence in the Gulf of Finland. Estonian Marine Institute, University of Tartu, Mäealuse 10a, 12618 Tallinn, Estonia, 48 p. (June 2005). PDF (1 MB)

Marchetti, Adrian. 2005. Ecophysiological aspects of iron nutrition and domoic acid production in oceanic and coastal diatoms of the genus Pseudo-nitzschia. Botany Department, University of British Columbia, Vancouver, Canada, 220 p. (April 2005). Abstract and link to PDF file

Salierno, James. 2005. Harmful algal bloom stressors alter behavior and brain activity in the killifish, Fundulus heteroclitus. University of Maryland, USA, 234 p. AbstractPDF (2.7 MB)


Boissonneault, Katie Rose. 2004. Gene discovery and expression profiling in the toxin-producing marine diatom, Pseudo-nitzschia multiseries (Hasle) Hasle. MIT/WHOI Joint Program on Biological Oceanography, Cambridge, MA, USA, 181 p. (September 2004). Abstract and link to PDF file

Fehling, Johanna. 2004. Diversity, ecology and domoic acid production of Pseudo-nitzschia spp. in Scottish waters. The Open University, Milton Keynes, UK, 309 p. (June 2004). Abstract

Koukaras, Konstantinos. 2004. Temporal and spatial distribution of DinophysisEhrenberg harmful blooms in Thermaikos Gulf (Greece). Department of Botany, School of Biology, Aristotle University of Thessaloniki, Thessaloniki, Greece, 156 p. (May 2004). PDF file of thesis summary (70 KB)

Leong, Sandric Chee Yew. 2004. Response of the dinoflagellate Alexandrium tamarense to nitrogen sources and concentrations: physiological, toxic and optical characteristics. Soka University, Japan, 209 p. DIALOG Abstract


Band Schmidt, Christine Johanna. 2003. Ciclo de vida, crecimiento y toxicidad de los dinoflagelados subtropicales: Alexandrium affine y Gymnodinium catenatum. Centro de Investigaciones Biológicas del Noroeste, La Paz, Mexico. (July 2004). Website for CIBNOR theses

Burgess, Vanessa Anne. 2003. Toxicology investigations with the Pectenotoxin-2 Seco Acids. Griffith University, Brisbane, Australia (February 2003). Abstract and link to PDF files

Dyble, Julianne. 2003. Detection and characterization of nitrogen-fixing cyanobacterial harmful algal bloom communities using molecular approaches. University of North Carolina at Chapel Hill, USA, 188 pp. DIALOG Abstract

Popels, Linda C. 2003. Investigations into the distribution, dark survival, and detection of Aureococcus anophagefferens. University of Delaware, USA. 115 p. DIALOG Abstract

Silvagni, Paul A. 2003. Comparative pathology and diagnosis of domoic acid toxicity. PhD Thesis, University of California, Davis, 139 p.

Siopsis, Maria. 2003. An individual-based model for the toxic algae species Pseudo-nitzschia multiseries. University of Tennessee, Knoxville, TN, USA, 112 p. (May 2003). PDF (1 MB)


Ajuzie, Cyril C. 2002. Studies on the harmful algal bloom phenomenon: a first perception and monitoring in Nigerian coastal waters, and the effects of the DSP-producing dinoflagellate Prorocentrum lima (Ehrenberg) Dodge on other aquatic biota. Université Libre de Bruxelles, Belgium, 260 p. Abstract

Colin, Sean P. 2002. Determination and characterization of resistance by populations of the copepod Acartia hudsonica to the toxic dinoflagellate Alexandrium sp. University of Connecticut, USA, 168 pp. DIALOG Abstract

Cusack, Caroline K. 2002. The diatom Pseudo-nitzschia in Irish waters. National University of Ireland, Galway, 380 p.

Etheridge, Stacey M. 2002. Ecophysiology and optical detection of harmful algal blooms. University of Connecticut, USA, 182 p. Abstract and link to PDF file

Haque, Shahroz M. 2002. Physiological regulation of toxin production in harmful Raphidophytes. Kagoshima University, Japan, 108 p. DIALOG Abstract

Maneiro Estraviz, Isabel M. 2002. Effects of harmful algae blooms on zooplankton community and role of zooplankton as vector of toxins through the food web. Universidad de Vigo, Spain, 167 pp. DIALOG Abstract

O'Boyle, Shane M. 2002. Summer flora of shelf waters around western Ireland: Interactions between physical dynamics and phytoplankton and consequences for potentially harmful algal events. National University of Ireland, Galway, Ireland, 309 p. DIALOG Abstract


Bargu, Sibel. 2001. Examining the place and role of krill in domoic acid poisoning events in Monterey Bay, CA. PhD Thesis, University of California at Santa Cruz, U.S.A. 130 pp. DIALOG Abstractexcerpts of slides from presentation given at the DIALOG V Symposium, held Oct. 19-25, 2004

Persich, Graziela R. 2001. Growth conditions, genetic identification and toxicity of the dinoflagellate Alexandrium tamarense (Lebour) Balech from southern Brazil. Universidade de Rio Grande at Rio Grande, Brazil, 100 p. DIALOG Abstract

Sengco, Mario R. 2001. The aggregation of clay minerals and marine microalgal cells: Physicochemical theory and implications for controlling harmful algal blooms. Massachusetts Institute of Technology, USA, 237 p. DIALOG Abstract

Vila, Magda. 2001. Harmful algal blooms in the Catalan coast (NW Mediterranean). Universitat de Barcelona (Spain), 179 pp. DIALOG Abstract


Lundholm, Nina. 2000. A study of Pseudo-nitzschia based on morphology and molecular phylogeny – with implications for the systematics of Bacillariaceae. University of Copenhagen

Penta, Bradley. 2000. Phytoplankton competition on the West Florida shelf: A simulation analysis with red tide implications. University of South Florida, USA, 194 p. DIALOG Abstract

Yang, Zhenbo B. 2000. Harmful algal blooms in selected Hong Kong coastal waters. The University of Hong Kong


Garcés, Esther. 1998. Proliferation of dinoflagellates along the Catalan Coast (Spain). PhD thesis (in Spanish). Website for thesis

Windust, Anthony J. 1998. The physiological ecology of diarrhetic shellfish poisoning (DSP) toxin production by the dinoflagellate Ecuviaella lima (Ehr.) Bütschli. Dalhousie University, Halifax, Nova Scotia, Canada. 214 p. [Chapter 5 discusses the growth of P. multiseries and P. pungens in a turbidostat culture system: "Simplex optimization of the growth rate of two species of Pseudo-nitzschia, with respect to irradiance and temperature, using a computer-controlled turbidostat culture system"] (PDF file of thesis, but without the figs; 251 KB)


Harper, Fiona Morag. 1997. Uptake, retention and elimination of cysts of the toxic dinoflagellate Alexandrium spp. by the blue mussel, Mytilus edulis. M.Sc. Thesis, Memorial University of Newfoundland, 95 p. (Abstract) (PDF file; 11.2 MB)


Xi, D. 1996. Mechanisms of maitotoxin and domoic acid action to elevate cytosolic free calcium. Charleston, SC, Medical University of South Carolina


Pan, Youlian. 1994. Production of domoic acid, a neurotoxin, by the diatom Pseudonitzschia pungens f. multiseries Hasle under phosphate and silicate limitation. Dalhousie University, Halifax, Nova Scotia, Canada, 245 p. DIALOG Abstract